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Johnny Otis, the bandleader, pioneering producer, singer, songwriter and radio host who was at the forefront of the rhythm and blues movement, died at his home in the Los Angeles suburb of Altadena on Tuesday. He was 90.

Mr. Otis had been in poor health for several years, said his manager, Terry Gould.

Best known for his 1958 rock and roll spoof "Willie and the Hand Jive," Mr. Otis, born John Veliotes in 1921 in Vallejo, was the son of Greek immigrants who ran a grocery store in Berkeley. Growing up in an integrated neighborhood made a big impression on him.

"Black culture captured me," Mr. Otis said. "I loved it, and it was richer and more fulfilling and more natural. I thought it was mine."

Johnny Otis passes away

"When the snows fall and the white winds blow, the lone wolf dies but the pack survives."
Winter is Coming

Now this is the Law of the Jungle—as old and as true as the sky; And the wolf that shall keep it may prosper, but the wolf that shall break it must die.
As the creeper that girdles the tree-trunk, the Law runneth forward and back; For the strength of the Pack is the Wolf, and the strength of the Wolf is the Pack.